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Vauxia gracilenta









3D animation of Vauxia bellula and other sponges (Choia ridleyi, Diagoniella cyathiformis, Eiffelia globosa, Hazelia conferta, Pirania muricata, and Wapkia elongata) and Chancelloria eros a sponge-like form covered of star-shaped spines.
ANIMATION BY PHLESCH BUBBLE © ROYAL ONTARIO MUSEUM
3D model of Vauxia bellula.
ANIMATION BY PHLESCH BUBBLE © ROYAL ONTARIO MUSEUM
Vauxia bellula (ROM 53564). Highly branched specimen showing the stalk base. Specimen height = 65 mm. Specimen dry – polarized light. Collins Quarry on Fossil Ridge.
© ROYAL ONTARIO MUSEUM. PHOTO: JEAN-BERNARD CARON
Vauxia bellula (USNM 66508) – Lectotype. A simple non-branching specimen showing the typical endosomal pattern for this species. Specimen height = 75 mm. Specimen dry – direct light (left), wet – direct light (right). Walcott Quarry.
© SMITHSONIAN INSTITUTION – NATIONAL MUSEUM OF NATURAL HISTORY. PHOTOS: JEAN-BERNARD CARON
Vauxia densa (GSC 8231 counterpart of USNM 66517) – Lectotype. Fragment of a large and simple branching form. Specimen height = 152 mm. Specimen dry – direct light (left), dry – polarized light (right). Walcott Quarry.
© GEOLOGICAL SURVEY OF CANADA. PHOTOS: JEAN-BERNARD CARON
Vauxia gracilenta (USNM 66511) – Paralectotype. A small branching specimen with a conspicuous dermal layer obscuring the endosomal skeleton. Specimen height = 31 mm. Specimen wet – polarized light. Walcott Quarry.
© SMITHSONIAN INSTITUTION – NATIONAL MUSEUM OF NATURAL HISTORY. PHOTO: JEAN-BERNARD CARON
Vauxia irregulara (ROM 53572) – Holotype. Non branching form showing irregular skeletal structures typical of the species. Specimen length = 60 mm. Specimen wet – direct light (left), wet – polarized light (right). Walcott Quarry.
© ROYAL ONTARIO MUSEUM. PHOTOS: JEAN-BERNARD CARON
Vauxia gracilenta (USNM 66515) – Lectotype. A large branching specimen with a conspicuous dermal layer still covering parts of the endosomal skeleton. Specimen height = 74 mm. Specimen dry – polarized light (left), wet – polarized light (middle and right). Walcott Quarry.
© SMITHSONIAN INSTITUTION – NATIONAL MUSEUM OF NATURAL HISTORY. PHOTOS: JEAN-BERNARD CARON
Vauxia venata (USNM 66520) – Lectotype. Fragment of a specimen with broad branches typical of the species. Specimen length = 94 mm. Specimen dry – direct light (left), wet – polarized light (right). Walcott Quarry.
© SMITHSONIAN INSTITUTION – NATIONAL MUSEUM OF NATURAL HISTORY. PHOTOS: JEAN-BERNARD CARON
Taxonomy:
Vauxia was placed within the hexactinellids by Walcott in his 1920 original description but Rigby (1980) transferred the genus and family to the Demospongea. Demosponges, the same group that are harvested as bath sponges, represent the largest class of sponges today.
Vauxia – from Mount Vaux (3,319 m), a mountain Peak in Yoho National Park, British Columbia. The name refers to William Sandys Wright Vaux (1818-1885) an antiquarian at the British Museum.
gracilenta – from the Latin gracilis, “slender,” referring to the delicate structure of the sponge.
Burgess Shale and vicinity: V. bellula Walcott, 1920; V. densa Walcott, 1920; V. irregulara Rigby and Collins, 2004; V. venata Walcott, 1920.
Other deposits: none.
Age & Localities:
Burgess Shale and vicinity: Vauxia species are known in the Walcott, Raymond and Collins Quarries on Fossil Ridge, the Trilobite Beds, Tulip Beds (S7) and the Collins Quarry on Mount Stephen, and smaller sites on Mount Field and Odaray Mountain. Vauxia is also known from Monarch in Kootenay National Park.
Other deposits: V. bellula Walcott, 1920 from the Middle Cambrian Wheeler and Marjum Formations in Utah (Rigby et al., 2010); V. magna Rigby, 1980 from the Middle Cambrian Spence Shale in Utah (Rigby, 1980).
History of Research:
This sponge was originally described by Walcott in 1920. The genus was reviewed by Rigby (1980) and the species redescribed by Rigby (1986) and Rigby and Collins (2004) in their examination of the Burgess Shale sponges.
Description:
Specimens of Vauxia gracilenta can range from simple unbranched forms to more complex branching forms and reach up to 8 cm in height. Each branch is deeply conical and almost cylindrical, with a simple open central cavity (spongocoel) ending in a rounded of flat opening (osculum). The skeleton is double layered with a thin dermal layer and an inner layer (endosomal). The dermal layer has small openings (ostia) and is composed of a dense network of ladder-like fibers supported by radial fibers from the inner layer. The inner layer forms a regular reticulated net-like skeleton of fibers with 4-6 sided polygons which is characteristic of the genus and species. The fibrous elements (spongin) represent tough collagen proteins. There is no evidence of siliceous spicules in the skeleton.
The different species have been identified mostly based on variations of the skeletal elements and the shape of the branches. Some species can reach up to at least 15 cm in height (V. bellula, V. densa).
Vauxia is relatively common in the Raymond Quarry and other sites on Mount Stephen but is rare in the Walcott Quarry where it represents less than 0.05% of the community (Caron and Jackson, 2008).
Ecology:
Vauxia would have lived attached to the sea floor. Particles of organic matter were extracted from the water as they passed through canals in the sponge’s wall.
References:
CARON, J.-B. AND D. A. JACKSON. 2008. Paleoecology of the Greater Phyllopod Bed community, Burgess Shale. Palaeogeography, Palaeoclimatology, Palaeoecology, 258: 222-256.
RIGBY, J. K. 1980. The new Middle Cambrian sponge Vauxia magna from the Spence Shale of Northern Utah and taxonomic position of the Vauxiidae. Journal of Paleontology, 54(1): 234-240.
RIGBY, J. K. 1986. Sponges of the Burgess Shale (Middle Cambrian), British Columbia. Palaeontographica Canadiana, 2: 1-105 p.
RIGBY, J. K. AND D. COLLINS. 2004. Sponges of the Middle Cambrian Burgess Shale and Stephen Formations, British Columbia. Royal Ontario Museum Contributions in Science (1): 155 p.
RIGBY, J. K., S. B. CHURCH AND N. K. ANDERSON. 2010. Middle Cambrian Sponges from the Drum Mountains and House Range in Western Utah. Journal of Paleontology, 84: 66-78.
WALCOTT, C. D. 1920. Middle Cambrian Spongiae. Cambrian Geology and Paleontology IV. Smithsonian Miscellaneous Collections, 67(6): 261-365.
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