The Burgess Shale

Cambroraster falcatus

A sediment sifter with a spaceship-shaped carapace

Cambroraster falcatus, isolated H-element ROMIP 65316


Kingdom: Animalia
Phylum: Arthropoda
Higher Taxonomic assignment: Order Radiodonta, Family Hurdiidae
Species name: Cambroraster falcatus

With its single pair of jointed frontal appendages, lateral swimming flaps, and circular mouth structure, Cambroraster possesses all the hallmarks of Radiodonta, part of the stem group to the true arthropods which also includes the iconic Anomalocaris (Collins 1996). The frontal appendages with comb or rake-like inner spines are characteristic of the radiodont family Hurdiidae. Phylogenetic analysis found it to be closely related to Titanokorys from the Burgess Shale and Zhenghecaris from the Chengjiang deposit, which share similarities in carapace shape and a large number of finely-spaced spines on the appendages (Caron and Moysiuk 2021).

Described by: Moysiuk and Caron
Description date: 2019

CambrorasterCambro, for Cambrian; raster, for the rake-like morphology of the inner spines on the frontal appendages.

falcatus – meaning sickle-shaped, but more specifically in reference to the dorsal carapace’s resemblance to the fictional Millennium Falcon starship in the Star Wars franchise.

Type Specimens: Holotype ROMIP 65078; Paratypes ROMIP 65079, 65081, 65083, 65084, 65092, at the Royal Ontario Museum.
Other species:

Burgess Shale and vicinity: None

Other deposits: Cambroraster sp. from the early Cambrian Chengjiang biota (Liu et al. 2020); Cambroraster cf. C. falcatus from the mid-Cambrian Mantou Formation of north China (Sun et al. 2020).

Age & Localities:

Middle Cambrian, Wuliuan stage, Burgess Shale Formation (around 507 million years old).
Principal localities:

Marble Canyon and Mount Whymper / Tokumm Creek, Kootenay National Park, British Columbia.

History of Research:

Brief history of research:

Several specimens of Cambroraster were discovered at the Marble Canyon and North Tokumm sites in Kootenay National Park in 2014. Because of their distinctive shape, the head carapaces were nicknamed the “spaceship.” Isolated frontal appendages were initially tentatively assigned to the genus Hurdia (Caron et al. 2014). The affinities of Cambroraster were not well-understood until further finds of abundant material at North Tokumm in 2018. The genus and species were formally described in 2019 (Moysiuk and Caron 2019). 3D digital modeling of an appendage of Cambroraster found it to have the lowest potential degree of appendage articulation of any of the studied radiodontans (de Vivo et al. 2021).



The defining feature of Cambroraster falcatus is its large, horseshoe-shaped dorsal carapace. This carapace is rounded frontally and projects along the rear sides into elongate wing-like projections lined along their margins with small spines. The rear central part of the carapace extends into a bilobate projection. Between the lateral “wings” and central projection are deep notches that accommodate the elliptical eyes, which are directed upwards. On the underside, the head is protected by two additional plates, shaped like elongate paddles and joined together at the front by their narrow ends. A circular, tooth-lined jaw and a pair of jointed frontal appendages with five long, curving, strong rake-like inner spines are located on the underside, near the front of the head. The body is stout, shorter than the dorsal carapace, and composed of 11 segments bearing rows of stacked gill blades and short lateral swimming flaps plus four short tail blades.


Cambroraster is abundant in Kootenay National Park, being known from over 100 specimens. It is particularly abundant around the North Tokumm locality, and may occur by the dozens on certain bedding planes, suggesting gregarious mass moulting behaviour. Rarer remains are known from Marble Canyon and single, isolated carapace fragments are known from Mount Stephen and Mount Field.

Maximum Size:
About 300 mm


Life habits: Mobile, Nektobenthic
Feeding strategies: Carnivorous
Ecological Interpretations:

Like other hurdiids, Cambroraster shows adaptations to sweep feeding. Specifically, the stout and rigid frontal appendages are ill-suited to grasping large, mobile prey (de Vivo et al. 2021). Instead, the rake-like inner spines on the appendages form a rigid, basket-like apparatus of spines surrounding the mouth. Sideways movements of the appendages could have disturbed the sediment, sifting out burrowing organisms, and transferring them to the mouth for further processing (Moysiuk and Caron 2019). Compared to related hurdiids like Hurdia and Stanleycaris, the particularly numerous and finely-spaced, strong, hooked secondary spines on the inner spines could have enabled capture of minute benthic organisms, although larger prey may also have been consumed. As one of the largest animals in the Marble Canyon and Tokumm communities, Cambroraster would have been near the top of the food chain. The broad dorsal carapace, upward facing eyes, and stubby body suggest it spent most of its time near the sea floor (Moysiuk and Caron 2019). As in other radiodontans, swimming was facilitated by undulation of the lateral flaps while respiration would have been accomplished primarily through the rows of gill blades on the body (Usami 2006; Daley et al. 2013).


  • CARON, J.-B. and MOYSIUK, J. 2021. A giant nektobenthic radiodont from the Burgess Shale and the significance of hurdiid carapace diversity. Royal Society Open Science, 8: 210664.
  • CARON, J.-B., GAINES, R. R., ARIA, C., MÁNGANO, M. G. and STRENG, M. 2014. A new phyllopod bed-like assemblage from the Burgess Shale of the Canadian Rockies. Nature communications, 5: 1–6.
  • COLLINS, D. 1996. The “evolution” of Anomalocaris and its classification in the arthropod class Dinocarida (nov.) and order Radiodonta (nov.). Journal of Paleontology, 70: 280–293.
  • DALEY, A. C., BUDD, G. E. and CARON, J.-B. 2013. Morphology and systematics of the anomalocaridid arthropod Hurdia from the Middle Cambrian of British Columbia and Utah. Journal of Systematic Palaeontology, 11: 743–787.
  • LIU, Y., LEROSEY-AUBRIL, R., AUDO, D., ZHAI, D., MAI, H. and ORTEGA-HERNÁNDEZ, J. 2020. Occurrence of the eudemersal radiodont Cambroraster in the early Cambrian Chengjiang Lagerstätte and the diversity of hurdiid ecomorphotypes. Geological Magazine, 157: 1200–1206.
  • MOYSIUK, J. and CARON, J.-B. 2019. A new hurdiid radiodont from the Burgess Shale evinces the exploitation of Cambrian infaunal food sources. Proceedings of the Royal Society B, 286: 20191079.
  • SUN, Z., ZENG, H. and ZHAO, F. 2020. Occurrence of the hurdiid radiodont Cambroraster in the middle Cambrian (Wuliuan) Mantou Formation of North China. Journal of Paleontology, 94: 881–886.
  • USAMI, Y. 2006. Theoretical study on the body form and swimming pattern of Anomalocaris based on hydrodynamic simulation. Journal of Theoretical Biology, 238: 11–17.
  • DE VIVO, G., LAUTENSCHLAGER, S. and VINTHER, J. 2021. Three-dimensional modelling, disparity and ecology of the first Cambrian apex predators. Proceedings of the Royal Society B, 288.
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